Opponent has just cast his second creature spell in a turn with a vengevine in the graveyard.
Can I activate relic of progenitus while vengevine's ability is on the stack removing it from the game?
Why would you doubt that you can?
Whenever you cast a spell, if it's the second creature spell you cast this turn, you may return Vengevine from your graveyard to the battlefield.
“It's possible. But it involves... {checks archives} Nature's Revolt, Opalescence, two Unstable Shapeshifters (one of which started as a Doppelganger), a Tide, an animated land, a creature with Fading, a Silver Wyvern, some way to get a creature into play in response to stuff, some way to get a land into play in response to stuff (a different land from the animated land), and one heck of a Rube Goldberg timing diagram.”
-David DeLaney
Yep. Since vengevine has a triggered ability, the ability is what is on the stack, not the creature itself. This is what lets graveyard removal get rid of the creature.
Sidenote: Vengevine only triggers on the second creature, so they can't cast another at instant speed to dodge your graveyard removal.
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